|Abstract or Summary
- Classification based on metathoracic wing venation does not
coincide with the existing higher classification of the family Scutelleridae.
The wings of the genera in the Scutellerinae possess a
similar general pattern of venation which is quite distinct from that
of Eurygasterinae, Odontoscelinae, Odontotarsinae, and Pachycorinae.
The Scutellerinae wings possess three additional characters
not found in the other subfamilies: a second secondary vein (present
in all of the genera); an antevannal vein (present in three-fourths of
the genera), and a Pcu stridulitrum (present in one-half of the
genera). Based upon wing venation the genera at present included
in the Scutellerinae do not fit into the tribal classification.
The wings of the Scutellerinae fell into two natural groups,
those with the Pcu stridulitrum and those without it. Those without
the Pcu stridulitrum are more generalized than those with it.
The four other subfamilies in the Scutelleridae, Eurygasterinae,
Odontoscelinae, Odontotarsinae and Pachycorinae, cannot be
separated from one another on the basis of the characters associated
with the metathoracic wing. However, genera in these taxa could be
separated from each other in most cases.
The Pachycorinae are very homogeneous and very generalized
as a group. Two-thirds of the genera are generalized. The Odontoscelinae
are more heterogeneous and specialized than the Odontotarsinae
which are either generalized or intermediate-generalized.
Eurygaster Laporte and Xerobia Stal[superscript o] are included in the Eurygasterinae.
Eurygaster is slightly more generalized than Xerobia.
The taxonomic position of Tectocoris Hahn is uncertain. It
possesses an intermediate-generalized wing. The genus Macro-carenus Stal[superscript o] should be removed from the Scutelleridae because it has
a pentatomid type of metathoracic wing and the scutellum possesses a
frena. On the basis of these characters it belongs in the Pentatomidae.
In the subfamily Scutellerinae, a stridulitrum situated on the
underside of the wings at the base of the Pcu was observed for the
first time. It was present in 12 out of the available 24 genera. The
plectrum is located on the anterior-dorsal part of the abdomen and
takes the form of a heavily sclerotized, file-like schlerite.
Scutelleridae wings are more generalized than Pentatomidae
wings. Selected examples of other families within the Pentatomoidea
were examined. Cyrtocoris White has the most generalized wing. Tessartoma Berthold has an intermediate metathoracic wing. The
Corimelaenidae Thyreocoris scarabaeoides (L.) and Corimelaena
nigra Dallas have very specialized wings. The wing of the Cydnidae,
Sehirus cinctus albonotatus Dallas is intermediate. Brachyplatys
vahlii (F.) (Plataspidae) and Phloea subquadrata Spinola have intermediate
The interpretation of Davis (1966) on metathoracic wing nomenclature
has been used in this paper. The higher classification follows
the classification used by Stal[superscipt o] (1873) with minor modifications.
There are many useful taxonomic characters associated with
the metathoracic wing. The taxonomic level at which these characters
are useful varies from group to group. Metathoracic wing
venation proved to be a very useful structure in classification at the
generic levels. On the other hand, at the higher taxonomic levels it
is possible to define only the subfamily Scutellerinae. It is necessary
to use other characters in conjunction with wing characters to
define the Eurygasterinae, Odontoscelinae, Odontotarsinae and