• A technique for measuring in vitro respiration was investigated to understand why rates were higher than those reported in vivo and to elucidate trends within mature Pseudotsuga menziesii (Douglas‐fir) trees.
• Extracted increment cores were divided into 3–4 radial depths and a gas chromatograph was used to compare respiration...
Live western larch, Larix occidentalis Nutt., a tree species resistant to the Douglas-fir beetle, Dendroctonus pseudotsugae Hopkins, produces the monoterpene 3-carene in higher concentrations compared to Douglas-fir, the preferred host of D. pseudotsugae (Reed et al. 1986). The inhibitory effects on attraction to aggregation pheromones and toxicity of 3-carene to...
The Chugach National Forest has been using prescribed fire as a wildlife habitat management tool since l977. Between 1977 and 1997 about 4,000 hectares have been burned on the Kenai Peninsula to promote regeneration of woody plant species used by moose (Alces alces). Browse species include paper birch (Betula papyrifera),...
The MODIS NBAR (M0D43B4) data space is explored in terms of biophysical variables with the objective of formulating a MODIS transformation relevant to global vegetation studies. The basic ideas of transform formulation were borrowed from the development of the TM Tasseled Cap transformation, but with differences in sampling strategy and...
During the last century, fire suppression, grazing, and climate change have caused sagebrush grasslands to be altered in both function and form; juniper and sagebrush dominate the landscape at the expense of herbaceous plants. Management efforts to reduce juniper and sagebrush overstory in order to enhance herbaceous components of the...
This paper was published in: Deal, R.L. and C.A. Harrington, eds. 2006. Red alder—a state of knowledge. General Technical Report PNW-GTR-669. Portland, OR: U.S. Department of Agriculture, Pacific Northwest Research Station. 150 p.
To increase our understanding of the timber harvesting behavior of family forest owners, I developed theoretical and empirical timber harvesting models for family forest owners in the southeastern United States. An individual-choice model was the basis for the models. Family forest owners were modeled as utility-maximizers who made harvesting decisions...
Stream systems in the Pacific Northwest have come to be understood in the absence of beaver (Castor canadensis). To understand the effect of beaver upon riparian plant communities, four basins in the lower Alsea drainage were examined to determine the effect of beaver and their impoundments on streamside herbaceous/shrub and...
Field studies were conducted from 1996 to 1998 in three villages in a Highland watershed in Mae Hong Son Province, northern Thailand. The objectives of the study were to classify and define the fruit-based agroforestry cropping system and investigate
management activities used by villagers. In the first part of the...
To effectively manage for biodiversity at broad, ecosystem scales, the influences of habitat structure at multiple spatial scales on vertebrate species must be understood. There are few studies on the broad-scale habitat requirements of stream amphibians despite their importance in streams in forest ecosystems in the Pacific Northwest (PNW) as...
Thinning of young Douglas-fir forests has the potential to enhance structural diversity and improve habitat for wildlife. I examined the effects of thinning and thinning intensity on abundance and demographic characteristics of forest-floor small mammals in the Coast Range of Oregon 5 and 6 years after thinning had occurred. Thinning...
My objectives were to investigate the correlation of height and diameter growth in young Douglas-fir (Pseudotsuga menziesii) plantations with stand density, and to determine the effects of stand density on the canopy boundary layer conductance (g[subscript]ac) and microclimate characteristics of young Douglas-fir plantations. I measured annual height (h) and diameter...
Two experiments were conducted on the invasive plant species Potentilla recta in northeastern Oregon to investigate both biological and ecological characteristics of the species. Growth analysis was conducted on the plant at three sites over a period of two years (2002 and 2003). Relative growth rate (RGR) was calculated for...
The nomenclatural history of Gomphus sensu lato began with Persoon in
1797. Over 200 years species of Gomphus sensu lato have been variously classified
also under Cantharellus, Craterellus, Chloroneuron, Chlorophyllum,
Gloeocantharellus, Nevrophyllum, and Turbinellus.
Species of Gomphus sensu lato have been historically characterized as
having aboveground fruiting, fleshy basidiomata with...
The tree species and size structure of 9l old-growth forests dominated by Douglas-fir
in central western Oregon was characterized using complete inventories of all trees larger than 20cm dbh over a mean area of 17.1ha at each site. Douglas-fir accounted
for over 75% of the total average basal area (39.1...
Viability analysis of well-selected focal species can complement other types of conservation planning by revealing thresholds in habitat area and landscape connectivity that may not be evident from ecosystem-level evaluations. I used focal species analysis of five carnivores to suggest conservation planning guidelines for the Rocky Mountains and adjacent areas...
The Amazon Tropical Rain Forest is the largest tropical rain forest system, comprising approximately 65% of such forests on earth. Since the 1960's, human populations in the Brazilian Amazon have increased from two to over 20 million. Concomitant urbanization, forest conversion, and economic dependence on resource extraction have exerted severe...
The goal of this dissertation was to improve our understanding of age-related constraints on aboveground production of forest trees. Previous research suggesting that carbon uptake of old trees is hydraulically constrained by tree size was used as the
springboard for this research. Three specific working hypotheses were investigated: 1) compensation...
Prescribed burning is increasingly being used as a management tool to reduce potential fuels on the forest floor and promote stand vigor through removal of dense, crowded vegetation. Because fire affects both the standing vegetation and the amount of downed wood in the ecosystem, it is imperative to examine how...
Human activities are causing profound changes to the global environment, yet the potential consequences of these changes on rising atmospheric carbon dioxide and climate change are not well understood. Improving our understanding of these processes requires an enhanced awareness of how global ecosystems are changing and how these changes affect...
I conducted two separate studies, both related to the impacts of spring and fall prescribed fire on ponderosa pine (Pinus ponderosa Dougl. ex Loud.) forest soils in Eastern Oregon. The studies were either conducted at or linked to four stands of ponderosa pine, in the Malheur National Forest. Each stand...
Delayed tree mortality is an unpredictable occurrence when prescribed burning is implemented. Fire scorched trees may die as the result of crown scorching, stem charring, root injury, bark beetle attack, or through a combination of these factors. This study examined ponderosa pine mortality and the incidence of two bark beetle...
Hemlock woolly adelgid, Adelges tsugae Annand, is a serious threat to eastern hemlock, Tsuga canadensis Carriere, in the eastern United States. Infestations of A. tsugae are spreading rapidly and tree mortality caused by the insect is increasing yearly. As part of an integrated management approach, a classical biological control program...
Balancing resource extraction with protection of Pacific Northwest forest ecosystems requires understanding the population biology of ectomycorrhizal fungi. Design of landscape-scale management and research strategies requires understanding genet size, habitat requirements, and dispersal capabilities. The purpose of this dissertation was to gain knowledge about the distribution of genetic variation, systematics,...
A series of studies, in three western Oregon Douglas-fir plantations, was conducted to understand the physiological impacts of Swiss needle cast on Douglas-fir physiology. Four aspects of the disease complex were investigated: fungal colonization and assessment, plant-water relations, carbon assimilation and interaction with climate. Several techniques were developed and used...
Seasonal and annual patterns of N fluxes and concentrations in streamwater in six conifer-dominated watersheds at the H.J. Andrews Experimental Forest, OR, were studied to gain insight into the factors that influence N retention in this ecosystem. Processes affecting N flux in streamwater differed between organic (DON) and inorganic (DIN;...
I studied riparian forests of four western Oregon watersheds (dry south to wet north) to determine the multiscale controls on woody riparian vegetation. I conducted separate analyses of controls on plant distribution, diversity, and tree regeneration using vegetation and environmental data collected in two related field studies: (1) a multiscale...
A simple debris-slide model, employing a digital elevation model (DEM) and geological data, was used in a geographic information system (GIS) to map slope stability in the Andrews Experimental Forest, located in the western Cascade Range in Oregon, USA. To evaluate the contribution of error in elevation to the uncertainty...
A mechanistic simulation model of nitrogen fixation in dead wood was developed to help synthesize knowledge, develop hypotheses, and estimate rates of nitrogen fixation in the Pacific Northwest. In this model nitrogen fixation is directly controlled by log substrate, temperature, moisture, and oxygen content. Respiration and diffusion of
oxygen indirectly...
Hybrid poplar plantations (Populus trichocarpa x Populus deltoides) are a relatively new feature on the landscape in the Pacific Northwest of the United States, and these plantations may soon include genetically engineered trees. Meanwhile, many wild poplar populations (Populus spp.) are highly degraded due in part to logging, dams, grazing,...
Maritime cyclonic windstorms cause widespread disturbance to forested ecosystems in southeast Alaska. The consequence of this disturbance process on the movement, storage, and quality of soil carbon, forest hydrology and streamwater chemistry was studied along a windthrow disturbance sequence. Soil profiles were described and the thickness of the major organic...
This study characterized the nature and dynamics of interference in mixed red alder
(Alnus rubra Bong.)/Douglas-fir (Pseudotsuga menziesii [Mirb.] Franco) stands in the Pacific Northwest, USA. Long-term spatial and tree measurements from the Cascade Head (CH) and H.J. Andrews (HJA) Experimental Forests in western Oregon and Delezene Creek (DC), Washington...
Application of N fertilizer is a common forest management practice in the Pacific
Northwest, yet the long-term influence of fertilization on forest soil properties is not well known. Although elevated N often increases mineralization of C and N from labile organic matter, negative effects have been documented in recalcitrant organic...
A series of experiments were established to gain a better understanding of the extent to which Douglas-fir seeding root architecture may be manipulated and subsequent influences on seedling morphological and physiological development. The incorporation of amendments into nursery soils changed root architecture to some degree, but did not produce large...
The growth potential of Douglas-fir, grand fir and western hemlock advance regeneration in the forest understory of Douglas-fir-dominated forests was investigated to detennine the feasibility of selection cutting systems in northwest Oregon. This study compared the growth of Douglas-fir, grand fir and western hemlock advance regeneration along a light gradient...
Wood density, moisture content, tracheid width and cell wall size were examined in trees from plots that were sprayed for 5 years with chlorothalonil (Bravo®) fungicide to reduce the impact of Swiss needle cast (SNC) and from trees in adjacent unsprayed plots. The unsprayed (more heavily diseased) trees had significantly...
Beginning in the 1980s, large-scale commercial harvest of Tricholoma magnivelare created a need for management of forests to ensure sustainability, but little was known of the biology or ecology of this species to guide management decisions. Four of the five studies presented here explore the basic synecology of T. magnivelare...
The spruce weevil (Pissodes strobi) is a serious pest of Sitka spruce in Oregon. Weevils cause damage by killing the leader of a tree, resulting in defects such as crooks and forks that can render the tree unmerchantable. In this study, spruce stands 16-25 years old were surveyed for weevil...
Silviculture systems that involve commercial thinning may provide higher quality wildlife habitat than traditional clearcut systems, yet such systems have not been vigorously tested. This prospective study examined forest floor animal abundance and habitat relationships under three different silviculture conditions: clearcuts, commercial thins, and uncuts. Eighteen stands, six per treatment,...
These studies are part of the Demonstration of Ecosystem Management Options (DEMO) study, a program researching the effects of different levels and patterns of green-tree retention on ecological, economic, and social phenomena. We restricted our studies to the 15% basal area, evenly dispersed retention treatment. Our objectives were, first, to...
Simulations of stream-subsurface water exchange (hyporheic exchange) using a three-dimensional steady state groundwater flow model and a particle tracking model in unconstrained and constrained reaches of small (2nd-order) and intermediate (5th-order) mountain streams were conducted to estimate the effects of geomorphic features on the extent, volume and residence time of...
Arthropods are important food resources for birds. Forest management activities can influence shrub-dwelling arthropods by affecting the structure and composition of understory shrub communities. Changes in abundance and species composition of arthropod communities in turn may influence the distribution and abundance of insectivorous birds. I examined relationships among bird abundance,...
Adaptive ecosystem management is a new paradigm for managing federal forests which requires regular monitoring of ecosystem function and diversity to measure the effects of management. Managers need new strategies and tools to help them assess their progress in maintaining healthy, productive and biologically diverse forests. Biomonitoring of select forest...
Weed control in Pacific Northwest forests has been criticized for its potential impact on biodiversity. Changes were evaluated in conifer growth, diversity of vegetation in situ and of recruiting plants through different regeneration mechanisms after applying temporally-varied weed control by herbicide treatments based on the critical period concept. Conifer size...
The overall purpose of this study was to examine the root and shoot development of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) seedlings at two distinct time periods in seedling establishment (after I and 3 growing seasons) in response to fertilizer, stock size, vegetation control, and soil moisture treatments. Two separate experiments...
Thinning has the potential to increase structural diversity of managed forests for wildlife. During 1994-1996, I conducted experimental and observational studies using pitfall trapping to assess short-term and potential long-term effects of thinning on abundance and reproduction of forest-floor vertebrates in Douglas-fir (Pseudotsuga menzeisii) forests of the Oregon Coast Range....
A series of studies and replicated field sites were implemented in the Oregon Coast Range within Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) sapling plantations infected with varying levels of Swiss needle cast (SNC) caused by the fungus, Phaeocryptopus gaeumannii (Rhode) Petrak (PG). This research was conducted to understand the effects of...
To study the combined effects of seedling quality and fertilization at the time of planting, three experiments were established in western Oregon during the winter of 2000. The first experiment investigated the effect of preplanting rootvolume and fertilization rate on the field performance of 1+1 Douglas-fir seedlings during two growing...
Effects of small-scale disturbances on fecundity and growth rates of Linnaea
borealis L., Tiarella trifoliata L. var. unifoliata, and Trillium ovatum Pursh. were studied in
old-growth forests in the Tsuga heterophylla Zone on the west slope of the Cascade
Mountain Range of Oregon. Performance of each species was compared in...
Soil physical, chemical, and biological components as well as climate and physiographic characteristics can interact to have a great effect on forest regeneration and seedling growth response to different establishment activities. The objective of this project was to increase the understanding of the interactions between soil type, controlled-release fertilizers, and...
Effects of partial understory removal on belowground and aboveground ecosystem properties were investigated in 5 and 15 yr old Douglas-fir (Pseudotsuga menziesll (Mirb.) Franco) experimental plantations in western Oregon. Seedling survival was not affected by understory removal at age 5 yr.
But through age 15 yr, understory removal increased seedling...
Craterellus tubaeformis is a small to medium-sized forest mushroom that is fairly common in the Douglas-fir/western hemlock forests of the Pacific Northwestern United States and is most often associated with decayed coarse woody debris. In this study, the mycorrhizae of Craterellus tubaeformis in western Oregon is identified by DNA analysis...
The relationship between the level of Phaeocryptopus gaeumannii (Rohde) Petrak colonization and severity of Swiss needle cast (SNC) symptoms, the possibility of early testing of Douglas-fir (Pseudotsuga menziesii var. menziesii [Mirb.] Franco) for SNC tolerance, and geographic variation in coastal Oregon with respect to SNC tolerance were investigated. Comparisons between...
I evaluated fire occurrence, growth and recruitment and determined the fire history of 21 old and 20 young 8ha stands in Cascade, Siskiyou and mid-Coast mixed conifer and evergreen forests in southwestern Oregon. The rates and patterns of growth were measured and analyzed on 1,079 old-growth and 2,111
young stand...
Gene flow is a major evolutionary force and an important factor in the breeding and conservation of forest trees. I studied the applicability of SSR markers for measuring pollen-mediated gene flow (i.e., pollen flow) in Douglas-fir (Pseudotsuga menziesii [Mirb.] Franco). I developed SSR markers, tested alternative approaches for measuring pollen...
This study examined debris flows occurring in a 125 km² study area in the Blue River watershed in the western Cascade Mountains of Oregon over a 50-year period. Debris flow occurrence was found to be concentrated in a distinct zone of high activity occupying approximately half of the study area,...
This study aims to compare different methods of obtaining maximum growing season leaf area index (LAI) maps using remote sensing data, LAI and tree cover field data in a boreal forest near Thompson, Manitoba, Canada. The comparison includes aspatial methods such as traditional regression, inverse regression and reduced major axis,...
As concern over global warming intensifies, sequestration and storage of atmospheric CO2 has become an important scientific and policy issue. Confusion persists, however, over interpretation of forest carbon (C) source-sink dynamics, in part because conclusions drawn depend on temporal and spatial scales of analysis (e.g. day-week
scale vs. successional-scale), type...
This study assessed the effects of thinning on arthropod communities on understory plants in the Central Oregon Coast Range. Arthropods were sampled from five understory plants in five pairs of thinned and unthinned, young (50-80 yrs), managed Douglas-fir stands, from late May to mid-July of 1998. Vine maple (Acer circinatum),...
Agroforestry is a traditional farming practice in American Samoa that has helped to sustain the livelihood of the native population for centuries. These once self-sufficient islands have become economically dependent on U.S monies and other external resources during the past century. Dependency has caused a shift in the carrying capacity...
Use of Granier-style heat dissipation sensors to measure sap flow is common in plant physiology, ecology and hydrology. There has been concern that any change to the original Granier design invalidates the empirical relationship between sap flux density and the temperature difference between the probes. Here, we compared daily water...